Dr. Millhollen studied ringtailed lemur visual and olfactory communication as a graduate student at the Oregon National Primate Research Center. After discovering that lemurs could discriminate between the scents from different individuals, she went to Berenty Reserve, Madagascar, as a Duke University post-doctoral student. Anne found that both the ringtailed lemur and sifaka troops in Berenty’s rich gallery forest used scent marks to demarcate territorial boundaries. That research evolved into continuing studies about how food resources influence ringtailed lemur ranging, territorial behavior and population, how lemur foraging behavior affects the health of the forest, and how this interaction may determine the survival of gallery forest ecosystems in the south of Madagascar.