B.S., University of Washington, 1965. Ph.D., University of California, Berkeley, 1969 (Dan Koshland). Postdoctoral: Purdue University, 1970-73 (W. J. Ray, Jr.); University of Oregon, 1973-74 (F. W. Dahlquist). Honors and Awards: NIH Postdoctoral Fellow, 1971-72, Ersted Award for Distinguished Teaching, 1987. Mortar Board Professor of the Month - March 1988, March 1994, February 1999, November 1999, and October 2000. At Oregon since 1978.